A subset of these neurons is thought to be involved in using the optic flow to estimate the parameters of self-motion, such as yaw, roll, and sideward translation.
The problem arose as a result of insufficient sideward freedom of movement of the trailing axle when the locomotive traversed sharp curves and turnouts.
Some include a wide range of responses such as when cats arch their backs, erect their hairs and adopt a sideward posture to communicate fear or aggression.