These are just the rules for dualising four-dimensional polychora, except that the usual finite method of reciprocation about a concentric hypersphere can run into problems.
Physiological activity in the human organism that concerns the consumption or transportation of nutrients is also connected with the reciprocation of various genetic variants.
To gain the maximum potential work from the engine a high reciprocation rate is required, typically 80% faster than a double-acting counterflow type engine.
