The female ranges up to 25 mm (1 inch) in length, is wingless and ground-dwelling, and exclusively hunts mole crickets, whereas all other species of tiphiids attack beetle larvae.
Males communicate with neighboring males through vibrational signals, and the songs they project to flying females are harmonic chirps, rather than the trills produced by most mole crickets.
The tympanum or ear is located in the front tibia in crickets, mole crickets, and katydids, and on the first abdominal segment in the grasshoppers and locusts.