Locomotion techniques used include leaping from tree to tree, walking on two or four limbs, knuckle-walking, and swinging between branches of trees (brachiation).
The wrist is held in a stable, locked position during the support phase of knuckle-walking by means of strongly exed interphalangeal joints, and extended metacarpophalangeal joints.
However, chimpanzees and gorillas are the closest living relatives to humans, and share anatomical features including a fused wrist bone which may also suggest knuckle-walking by human ancestors.
This is supported by the evidence that gorilla and chimpanzee differ in their knuckle-walking related wrist anatomy and in the biomechanics of their knuckle-walking.
Pre-bipedal locomotion is probably best characterized as a repertoire consisting of terrestrial knuckle-walking, arboreal climbing and occasional suspensory activities, not unlike that observed in chimpanzees today.