The activities of mangrove fauna in the intertidal mudflats develop micromorphological features that trap and hold sediments to create a substratum for mangrove seeds.
Such subsistence of many attributes in a common substratum is the correct apposition ("samndhikaranya"), rather than the mere apposition as propounded by the advaita school.
Initial formation of podosomes seems to be influenced by the structure and composition of the underlying substratum including the presence and distribution of specific ligands.
The juxtaposition of features are sometimes without explanations using the deposition laws, probably in relation with post-sedimentary strain phenomena, eventually linked to substratum collapse.